Hemione

Its head is large, with ears smaller than those of the hemione, the shoulders thick, especially in the male, the limbs robust and stubbier than those of the hemiones and the asses.

They presumably belong to hemiones (hemiones were actually recorded from the same level at Rana Gundhai) and, in spite of some optimistic statements to the contrary, it is sometimes impossible to distinguish horses fom hemiones on isolated teeth alone.

Another possible area of disagreement is that asses and hemiones are associated unequivocally in the morphological tree, but less definitely in the genetic tree: whether hemiones and African ('true') wild asses are more closely related to horses or whether they are most closely related to each other and form a separate lineage has yet to be resolved unequivocally by the DNA data, and note that in the morphological data they are united by only a single derived condition (Fig. 1.1).